Macroevolution
evolution above species level, origination, diversification, and extinction of species over time
Microevolution
evolution within populations, changes in allele frequencies (both adaptive and neutral changes)
Biogeography
study of distribution of species across space and time
- Greater species diversity in tropical areas (amazon in south america and tropical deciduous in africa and asia), more ocean structures = more diversity
Species richness patterns
increase from polar to tropics -> latitudinal species richness gradient
Peninsular effect
organisms unaware of other islands via tree height, do not leave original island
Panama has high diversity due to continent interchange and mountainous landscape (allopatric speciation/speciation)
Species-time hypothesis
communities diversify (new species) with time, temperate less rich than tropical due to younger age (only recently recovered from ice age)
Species-area hypothesis
large areas have more species than small areas due to being able to support larger pop. (more resistant to extinction) and range of habitats
Species-energy hypothesis
available energy determines species richness, increased solar + water = more plants, more plant production -> more herbivores -> more predators, parasites, and scavengers
tropics have
largest land area
Hummingbirds
originated in south america, ocean creates harder migration/dispersal routes, some did cross over via panama land bridge
Coral reefs
more so in pacific than atlantic due to origination and more hurricanes in atlantic
fauna
assemblage of different animals species that live together (in ecosystem, region, or whole planet)
flora
assemblage of diff. Plant specie the live together
dispersal
movement of populations from one region to another with limited or no return exchange
vicariance
formation of geographic barriers to dispersal that divide a once continuous (living together) population distinct phylogenetic signature.
Example: marsupials (only found in australia and the americas) evolved via dispersal and vicariance
Interplay between speciation and extinction determines diversity
D1 (diversity) + originations – extinctions = D2 (new diversity)
new species originating faster than old species become extinct
fauna change in diversity due to intrinsic properties from large-scale changes in climate or environment
Standing diversity
# of species (OTU) present in particular area at given time, origination and extinction rates determine this number
Three evolutionary faunas
1. Cambrian 2. Paleozoic 3. Modern
Tidbit
high number of beetles due to new species originating faster than old species become extinct
taxonomic diversity is
positively correlated with mean temperature
anagenesis
wholesale transformation of lineage from one form to another (alt to splitting lineage or speciation)
punctuated equilibrium
periods of stasis punctuated by periods of change (associated with speciation events) (metrarabdotos fit the pattern)
Gradualism
slow, gradual morphological changes over time (can incllude speciation event; traditional view)
incomplete fossil records
- Old species went through anagenesis (new lineage/morphology formed)
- New species branched off from old one, rapid evolution -> new stasis (no change)
adaptive radiation
when α (origination rate) eclipses Ω (extinction rate) [ex. silverswords in the hawaiian archipelago]
Key innovation
enables adaptive radiation, is a novel trait that allows subsequent radiation and success of a clade
Example: evolution of nectar spurs allowed columbine (flowers) clade to diversify rapidly (radiation)
the cambrian explosion arose long after
animals began to diversify
fossil record documents how
animal phyla emerged
ecology of ocean changed during
cambrian, giving rise to new species
cloudina- predator earliest signs
Background extinction
normal rate of extinction for taxa or biota
Mass extinction
statistically significant increase above background extinction rates
Example: asteroid 66 mya -> mass extinction, known via K-T boundary with high levels of rare iridium, impact crater near yucatan, and quartz crystals
Human emissions currently responsible for
imbalance in earth’s energy budget -> global warming, etc.
However, diversity is positively correlated with mean temperature
ocean acidification
is the decrease in the pH of the earth's ocean
average fell and primary cause are human carbon dioxide emissions
Biotic factors
habitat loss via deforestation (i.e. tigers) is biggest biotic factor, others are invasive species, pollution, and direct exploitation
Abiotic factors
carbon dioxide emission increase -> warmer temps
Leads to ocean acidification: decrease in ocean pH due to increasing CO2 levels and depletion of oxygen