Heritability
ability of a trait to be passed on to offspring
Evolution
Decent with modification
Adaptation
inherited characteristics of organisms that enhance their fitness in specific envrionments
Fitness
quantifiable measure of survival and reproductive success
Natural Selection
process in which individuals that have certain inherited traits tend to have a higher fitness because of those adaptations
For natural selection to work: (3 aspects must occur)
Variation: 1. must be present in the population
2. must affect fitness
3. must be inherited
Artificial selection
the process in which humans modify organisms by selecting and breeding organisms with certain traits
Homologous structures
structures in different species that have a general theme because of common ancestry, but structures may vary in form and function
Vestigial structures
remnants of structures that serve an important function in the ancestor, but not in the present organism
Biogeography
the geographic distribution of species
Convergent evolution
the independent evolution of similar features in different lineages due to similar environmental pressures
microevolution
The change in allele frequencies in a population over time
populations
all the individuals of one species in a particular area
Genes
a discrete unit of hereditary info. consisting of a specific nucleotide sequence in DNA
Genotype
the genetic makeup, or set of alleles, of an organism
Allele
any alternative versions of a gene that may produce distinguishable phenotypic effects
Phenotype
observable traits of an organism that are determined by genotype
Dominant phenotype
homozygous dominant or heterozygous genotype
Recessive phenotype
homozygous recessive genotype
genetic variation
differences among individuals in the composition of their genes or other DNA segments
Phenotypic plasticity
variations in appearance due to environmental influences
2 ways to measure genetic variation:
1. nucleotide variability- at the molecular level of DNA
2. gene variability- at the whole gene level, which is quantified by average heterozygosity
Geographic variation
differences in the genetic composition of separate populations
cline
a graded change in character along a geographic axis
Sources of genetic variation
1. Formation of new alleles through mutation
2. Altering gene number or position through chromosomal changes that delete, disrupt, or rearrange loci
3. Rapid reproduction can increase mutation rates
4. Sexual reproduction due to: crossing over, independent assortment of alleles, fertilization
Hardy-Weinberg principle: original proportions of genotypes within a population will remain the same if:
1. no mutations
2. no gene flow
3. random mating occurs
4. population size is large
5. no selection occurs
Hardy Weinberg allele frequency equation
p+q=1
Hardy Weinberg genotype frequency equation
p(squared)+ 2pq + q(squared)=1
Genetic drift
chance events that can cause unpredictable fluctuations in allele frequencies from one generation to the next
Genetic drift example: Founder effect
When a few individuals become isolated from a larger population
Genetic drift example: Bottleneck effect
a reduction of population size due to a sudden change in environment
Gene flow
movement of alleles into or out of a population
immigration- move into
emigration- move out of
Natural Selection
process in which individuals that have certain inherited traits tend to have higher fitness because of those adaptations
Relative fitness
contribution an individual makes to the gene pool of the next generation relative to the contributions of other individuals
Directional-
natural selecting can alter frequency distribution
one end of distribution is favored
Disruptive-
natural selecting can alter frequency distribution
both ends of the distribution are favored
Stabilizing-
natural selecting can alter frequency distribution
intermediates of the distribution are favored
Sexual selection
a form of selection in which individuals with certain phenotypes are most likely than others to obtain mates
Intra-sexual selection
individuals of the same sex compete for access to mates
Inter-sexual selection
individuals of one sex choose their mates of the opposite sex
Balancing selection
occurs when natural selection maintains variation in the population
2 ways: Heterozygote advantage
Frequency dependent selection
Heterozygotes advantage
heterozygotes have a greater fitness than both homozygotes
Frequency-dependent selection
the fitness of the phenotype depends on how common it is in the population
Why natural selection cannot fashion perfect organisms????
1. selection can act only on existing variations
2. evolution is limited by historical constraints
3. adaptations are often compromises
4. chance, natural selection, and the environment interact
Speciation
the process by which one species splits into 2 or more species
Biological species concept
defines species as a group of populations whose members have the potential to interbreed in nature and produce viable, fertile offspring, but do not produce viable, fertile offspring with member of other such groups.
Reproductive isolation
the existence of biological factors(barriers) that impede members of 2 species from interbreeding and producing viable, fertile offspring
pre-zygotic barrier
block fertilization from occuring
Post-zygotic barriers
reproductive isolation occurs after the hybrid zygote is formed
Habitat isolation
species occupy different habitats with in the same area and therefore rarley encounter each other
Temporal isolation
species that breed during different times of the day, season, or years
Behavioral isolation
courtship rituals that attracts mates and enables mate recognition
Mechanical isolation
Mating is attempted, but morphological differences prevent a successful completion
Gametic isolation
mating is attempted, but the sperm of one species is unable to fertilize the egg of another species
Reduced hybrid viability
Zygote forms but either the embryo does not develop or the offspring doesn't survive in its environment
Reduced hybrid fertility
the hybrid is healthy, but is infertile and cannot produce offspring
Hybrid breakdown
when a viable, fertile hybrid reproduces, but their offspring are feeble or sterile
Morphological species concept-
characterizes species by body shape and structure features
Ecological Species concept
defines species based on ecological niche
Phylogenetic species concept
defines species as the smallest group of individuals that share a common ancestor
Allopatric speciation
occurs because gene flow is interrupted when a population becomes geographically isolated
Sympatric speciation
speciation occurs without geographic separation
Polyploidy
an accident in cell division that results in extra set of chromosomes
Habitat differentiation
genetic factors enable a subpopulation to exploit a habitat or resource not used by the present population
Sexual selection
a subset of the population starts choosing the opposite sex based on specific characteristics
Hybrid zones
a region in which members of different species meet and mate, producing some offspring of mixed ancestry
Reinforcement
process of natural selection strengthening the pre-zygotic barriers reducing the chances of hybrid formation
Fusion
weak reproductive barriers increase gene flow between the 2 species causing the 2 hybrid species to fuse into single species
Stability
Hybrids continue to be produced because the hybrids survive and/or reproduce better than the parent species
Punctuated equilibrium
periods of stasis followed by sudden change
Gradualism
change occurs gradually over long periods of time
Macroevolution
broad pattern of evolution above the species level
-revealed through fossil record
4 main stages the could have produced the first sell:
1. the abiotic synthesis of small organic molecules, such as AA and nitrogenous bases
2. the joining of these small molecules into macromolecules
3. the packaging of these molecules into protocell droplets with membranes that maintained internal chemistry different from the surroundings
4. The origin of the self-replicating molecules that eventually made inheritance possible
Radiometric dating
a technique of dating rocks and fossils based on the fixed rate of radioactive isotope decay
Half life
Amount of time required for 50% of the parent isotope to decay
Eukaryotes, organisms with membrane-bound organelles, evolved through:
-infolding of plasma membrane--created ER, nuclear membrane
-endosymbiotic theory-cell engulfing a prokaryote---created mitochondria, chloroplasts, plastids
Why is Multicellularity important?
hypothesis?
Led to the great innovation of specialization
- Colonial unicellular organisms let to multicellular organisms
cambrian explosion
burst of evolutionary change that occured in a short geological period , which includes the appearance of many present day animal phyla
Continental drift
process in which the earth's mantle causes the continental plates to move
-promotes allopatric speciation on a grand scale
Adaptive radiation
periods of evolutionary change in which groups of organisms form many new species whose adaptations allow them to fill different ecological roles in their communities
Horse fossils show 3 trends:
1. larger size
2.more complex molar teeth
3. fewer toes
Permian period
most extensive loss. 96% of plants and animal species perished
Cretaceous period
most famous and well studies. Caused by meteorite causing mass forest fire and smoke blocked sun, dinosaurs died off while placental mammals flourished
Taxonomy
disciplin in which organisms are named and classified
Taxa
taxonomic unit at any level
Binomial nomenclature
the scientific name of an organism that consists of a genus and species
Systematics
a discipline focused on classifying organisms and determining their evolutionary relationships
Phylogeny
the evolutionary history of a species or group of species
phylogenetic tree
branching diagram that represent the evolutionary history of a group of organisms
nodes
represent common ancestor
branch point
where lineages diverge
polytomy
a branch point where more than 2 taxa emerge
sister taxa
groups that share an immediate common ancestor
Basal taxon
lineage that diverges early in the history of the group
Homology
genotypic and phenotypic similarities due to sharing ancestry
Analogy
Phenotypic similarities, but no shared ancestry
Convergent evolution
process of similar adaptations evolving in organisms from different evolutionary lineages due to similar environmental pressures
Outgroup
a taxa that is closely related to the group of organisms that you are studying, but are not part of your study group
synaphories
shared derived characters that represent departure from the ancestor
Sympleslomorphies
shared ancestral characters that evolved prior to the most recent common ancestor
Monophyletic
a group that consists of the ancestral species and all of its descendants
Paraphyletic
a group that consists of the ancestral species and some, but not all of its descendants
Polyphyletic
a group that doesn't include the closest common ancestor
3 domains?
Archaea
Bacteria
Eukarya
kingdoms of Eukarya:
Plantae
Fungi
Animalia
Protista